Effect of Continental Changes on the Distribution of Animals.It is impossible to exaggerate, or even adequately to conceive, the effect of these endless mutations on the animal world. Slowly but surely the whole population of living things must have been driven backward and forward from east to west, or from north to south, from one side of a continent or a hemisphere to the other. Owing to the remarkable continuity of all the land masses, animals and plants must have often been compelled to migrate into other continents, where in the struggle for existence under new conditions many would succumb; while such as were able to survive would constitute those wide-spread groups whose distribution often puzzles us. Owing to the repeated isolation of portions of continents for long periods, special forms of life would have time to be developed, which, when again brought into competition with the fauna from which they had been separated, would cause fresh struggles of ever increasing complexity, and thus lead to the development and preservation of every weapon, every habit, and every instinct, which could in any way conduce to the safety and preservation of the several species.
Changed Distribution proved by the Extinct Animals of Different Epochs.We thus find that, while the inorganic world has been in a state of continual though very gradual change, the species of the organic world have also been slowly changing in form and in the localities they inhabit; and the records of these changes and these migrations are everywhere to be found, in the actual distribution of the species no less than in the fossil remains which are preserved in the rocks. Everywhere the animals which have most recently become extinct resemble more or less closely those which now live in the same country; and where there are exceptions to the rule, we can generally trace them to some changed conditions which have led to the extinction of certain types. But when we go a little further back, to the late or middle Tertiary deposits, we almost always find, along with forms which might have been the ancestors of some now living, others which are only now found in remote regions and often in distinct continentsclear indications of those extensive migrations which have ever been going on. Every large island contains in its animal inhabitants a record of the period when it was last separated from the adjacent continent, while some portions of existing continents still show by the comparative poverty and speciality of their animals that at no distant epoch they were cut off by arms of the sea and formed islands. If the geological record were more perfect, or even if we had as good a knowledge of that record in all parts of the world as we have in Europe and North America, we could arrive at much more accurate results than we are able to do with our present very imperfect knowledge of extinct forms of life; but even with our present scanty information we are able to throw much light upon the past history of our globe and its inhabitants, and can sketch out with confidence many of the changes they must have undergone.
Summary of Evidence for the General Permanence of Continents and Oceans.As this question of the permanence of our continents or, rather, of the continental areas, lies at the root of all our inquiries into the past changes of the earth and its inhabitants, and as it is at present completely ignored by many writers, and even by naturalists of eminence, it will be well to summarise the various kinds of evidence which go to establish it.32 We know as a fact that all sedimentary deposits have been formed under water, but we also know that they were largely formed in lakes or inland seas, or near the coasts of continents or great islands, and that deposits uniform in character and more than 150 or 200 miles wide were rarely, if ever, formed at the same time. The further we go from the land the less rapidly deposition takes place, hence the great bulk of all the strata must have been formed near land. Some deposits are, it is true, continually forming in the midst of the great oceans, but these are chiefly organic and increase very slowly, and there is no proof that any part of the series of known geological formations exactly resembles them. Chalk, which is still believed to be such a deposit by many naturalists, has been shown, by its contained fossils, to be a comparatively shallow water formationthat is, one formed at a depth measured by hundreds rather than by thousands of fathoms. The nature of the formations composing all our continents also proves the continuity of those continents. Everywhere we find clearly marked shore and estuarine deposits, showing that every part of the existing land has in turn been on the sea-shore; and we also find in all periods lacustrine formations of considerable extent with remains of plants and land animals, proving the existence of continents or extensive lands, in which such lakes or estuaries could be formed. These lacustrine deposits can be traced back through every period, from the newer Tertiary to the Devonian and Cambrian, and in every continent which has been geologically explored; and thus complete the proof that our continents have been in existence under ever changing forms throughout the whole of that enormous lapse of time.
On the side of the oceans we have also a great weight of evidence in favour of their permanence and stability. In addition to their enormous depths and great extent, and the circumstance that the deposits now forming in them are distinct from anything found upon the land-surface, we have the extraordinary fact that the countless islands scattered over their whole area (with one or two exceptions only and those comparatively near to continental areas) never contain any Palæozoic or Secondary rocksthat is, have not preserved any fragments of the supposed ancient continents, nor of the deposits which must have resulted from their denudation during the whole period of their existence! The supposed exceptions are New Zealand and the Seychelles Islands, both situated near to continents and not really oceanic, leaving almost the whole of the vast areas of the Atlantic, Pacific, Indian, and Southern oceans, without a solitary relic of the great islands or continents supposed to have sunk beneath their waves.
CHAPTER VII
CHANGES OF CLIMATE WHICH HAVE INFLUENCED THE DISPERSAL OF ORGANISMS: THE GLACIAL EPOCH
Proofs of the Recent Occurrence of a Glacial EpochMorainesTravelled BlocksGlacial Deposits of Scotland: the "Till"Inferences from the Glacial Phenomena of ScotlandGlacial Phenomena of North AmericaEffects of the Glacial Epoch on Animal LifeWarm and Cold PeriodsPalæontological Evidence of Alternate Cold and Warm PeriodsEvidence of Interglacial Warm Periods on the Continent and in North AmericaMigrations and Extinctions of Organisms caused by the Glacial Epoch.
We have now to consider another set of physical revolutions which have profoundly affected the whole organic world. Besides the wonderful geological changes to which, as we have seen, all continents have been exposed, and which must, with extreme slowness, have brought about the greater features of the dispersal of animals and plants throughout the world, there has been also a long succession of climatal changes, which, though very slow and gradual when measured by centuries, may have sometimes been rapid as compared with the slow march of geological mutations.
These climatal changes may be divided into two classes, which have been thought to be the opposite phases of the same great phenomenoncold or even glacial epochs in the Temperate zones on the one hand, and mild or even warm periods extending into the Arctic regions on the other. The evidence for both these changes having occurred is conclusive; and as they must be taken account of whenever we endeavour to explain the past migrations and actual distribution of the animal world, a brief outline of the more important facts and of the conclusions they lead to must be here given.
We have now to consider another set of physical revolutions which have profoundly affected the whole organic world. Besides the wonderful geological changes to which, as we have seen, all continents have been exposed, and which must, with extreme slowness, have brought about the greater features of the dispersal of animals and plants throughout the world, there has been also a long succession of climatal changes, which, though very slow and gradual when measured by centuries, may have sometimes been rapid as compared with the slow march of geological mutations.
These climatal changes may be divided into two classes, which have been thought to be the opposite phases of the same great phenomenoncold or even glacial epochs in the Temperate zones on the one hand, and mild or even warm periods extending into the Arctic regions on the other. The evidence for both these changes having occurred is conclusive; and as they must be taken account of whenever we endeavour to explain the past migrations and actual distribution of the animal world, a brief outline of the more important facts and of the conclusions they lead to must be here given.
Proofs of the Recent Occurrence of a Glacial Epoch.The phenomena that prove the recent occurrence of glacial epochs in the temperate regions are exceedingly varied, and extend over very wide areas. It will be well therefore to state, first, what those facts are as exhibited in our own country, referring afterwards to similar phenomena in other parts of the world.
Perhaps the most striking of all the evidences of glaciation are the grooved, scratched, or striated rocks. These occur abundantly in Scotland, Cumberland, and North Wales, and no rational explanation of them has ever been given except that they were formed by glaciers. In many valleys, as, for instance, that of Llanberris in North Wales, hundreds of examples may be seen, consisting of deep grooves several inches wide, smaller furrows, and striæ of extreme fineness wherever the rock is of sufficiently close and hard texture to receive such marks. These grooves or scratches are often many yards long, they are found in the bed of the valley as well as high up on its sides, and they are almost all without exception in one general directionthat of the valley itself, even though the particular surface they are upon slopes in another direction. When the native covering of turf is cleared away from the rock the grooves and striæ are often found in great perfection, and there is reason to believe that such markings cover, or have once covered, a large part of the surface. Accompanying these markings we find another, hardly less curious phenomenon, the rounding off or planing down of the hardest rocks to a smooth undulating surface. Hard crystalline schists with their strata nearly vertical, and which one would expect to find exposing jagged edges, are found ground off to a perfectly smooth but never to a flat surface. These rounded surfaces are found not only on single rocks but over whole valleys and mountain sides, and form what are termed roches moutonnées, from their often having the appearance at a distance of sheep lying down.
Now these two phenomena are actually produced by existing glaciers, while there is no other known or even conceivable cause that could have produced them. Whenever the Swiss glaciers retreat a little, as they sometimes do, the rocks in the bed of the valley they have passed over are found to be rounded, grooved, and striated just as are those of Wales and Scotland. The two sets of phenomena are so exactly identical that no one who has ever compared them can doubt that they are due to the same causes. But we have further and even more convincing evidence. Glaciers produce many other effects besides these two, and whatever effects they produce in Switzerland, in Norway, or in Greenland, we find examples of similar effects having been produced in our own country. The most striking of these are moraines and travelled blocks.
Moraines.Almost every existing glacier carries down with it great masses of rock, stones, and earth, which fall on its surface from the precipices and mountain slopes which hem it in, or the rocky peaks which rise above it. As the glacier slowly moves downward, this débris forms long lines on each side, or on the centre whenever two glacier-streams unite, and is deposited at its termination in a huge mound called the terminal moraine. The decrease of a glacier may often be traced by successive old moraines across the valley up which it has retreated. When once seen and examined, these moraines can always be distinguished almost at a glance. Their position is most remarkable, having no apparent natural relation to the form of the valley or the surrounding slopes, so that they look like huge earthworks formed by man for purposes of defence. Their composition is equally peculiar, consisting of a mixture of earth and rocks of all sizes, usually without any arrangement, the rocks often being huge angular masses just as they had fallen from the surrounding precipices. Some of these rock masses often rest on the very top of the moraine in positions where no other natural force but that of ice could have placed them. Exactly similar mounds are found in the valleys of North Wales and Scotland, and always where the other evidences of ice-action occur abundantly.
A GLACIER WITH MORAINES.
Travelled Blocks.The phenomenon of travelled or perched blocks is also a common one in all glacier countries, marking out very clearly the former extent of the ice. When a glacier fills a lateral valley, its foot will sometimes cross over the main valley and abut against its opposite slope, and it will deposit there some portion of its terminal moraine. But in these circumstances the end of the glacier not being confined laterally will spread out, and the moraine matter will be distributed over a large surface, so that the only well-marked token of its presence will be the larger masses of rock that may have been brought down. Such blocks are found abundantly in many of the districts of our own country where other marks of glaciation exist, and they often rest on ridges or hillocks over which the ice has passed, these elevations consisting sometimes of loose material and sometimes of rock different from that of which the blocks are composed. These are called travelled blocks, and can almost always be traced to their source in one of the higher valleys from which the glacier descended. Some of the most remarkable examples of such travelled blocks are to be found on the southern slopes of the Jura. These consist of enormous angular blocks of granite, gneiss, and other crystalline rocks, quite foreign to the Jura mountains, but exactly agreeing with those of the Alpine range fifty miles away across the great central valley of Switzerland. One of the largest of these blocks is forty feet diameter, and is situated 900 feet above the level of the Lake of Neufchatel. These blocks have been proved by Swiss geologists to have been brought by the ancient glacier of the Rhone which was fed by the whole Alpine range from Mont Blanc to the Furka Pass. This glacier must have been many thousand feet thick at the mouth of the Rhone valley near the head of the Lake of Geneva, since it spread over the whole of the great valley of Switzerland, extending from Geneva to Neufchatel, Berne, and Soleure, and even on the flanks of the Jura, reached a maximum height of 2,015 feet above the valley. The numerous blocks scattered over the Jura for a distance of about a hundred miles vary considerably in the material of which they are composed, but they are found to be each traceable to a part of the Alps corresponding to their position, on the theory that they have been brought by a glacier spreading out from the Rhone valley. Thus, all the blocks situated to the east of a central point G (see map) can be traced to the eastern side of the Rhone valley (l e d), while those found towards Geneva have all come from the west side (p h). It is also very suggestive that the highest blocks on the Jura at G have come from the eastern shoulder of Mont Blanc in the direct line h B F G. Here the glacier would naturally preserve its greatest thickness, while as it spread out eastward and westward it would become thinner. We accordingly find that the travelled blocks on either side of the central point become lower and lower, till near Soleure and Geneva they are not more than 500 feet above the valley. The evidence is altogether so conclusive that, after personal examination of the district in company with eminent Swiss geologists, Sir Charles Lyell gave up the view he had first adoptedthat the blocks had been carried by floating ice during a period of submergenceas altogether untenable.33